Lycosidae Sundevall, 1833

Family |   ACCEPTED
N.B.: according to Brignoli, 1983c: 432, "Guy (1966, 1969) has proposed many synonymies and has downgraded many genera to subgenus rank; as this author based his conclusions more on nomenclature than on the study of material (or even of the original descriptions)", Platnick (WSC 15.0) did not accept Guy's conclusions; the changes by Guy, 1966 are noted (except where more recent work has superseded them); Guy, 1969 is merely a literature compilation that reflects those changes and is not cited separately; several genera of Lycosidae have been found not to be monophyletic (Murphy et al., 2006). Piacentini & Ramírez, 2019: 233-237 confirm ten valid subfamilies of Lycosidae: Venoniinae Lehtinen & Hippa, 1979, Zoicinae Lehtinen & Hippa, 1979 (= Piratinae Zyuzin, 1993), Evippinae Zyuzin, 1985, Sosippinae Dondale, 1986 (= Hygrolycosinae He & Song, 1996), Artoriinae Framenau, 2007, Tricassinae Alderweireldt & Jocqué, 1993 (= Arctosinae He & Song, 1996), Hippasinae Simon, 1898, Allocosinae Dondale, 1986, Pardosinae Simon, 1898 (= Wadicosinae Zyuzin, 1985) and Lycosinae Sundevall, 1833; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

In synonymy:

Boebe O. Pickard-Cambridge, 1885 = Evippa Simon, 1882 (Simon, 1898a: 362)

Evippella Strand, 1906 = Evippa Simon, 1882 (Alderweireldt, 1991: 360)

Gen. Evippa Simon, 1882 urn:lsid:nmbe.ch:spidergen:01615
N.B.: considered a senior synonym of Boebe O. Pickard-Cambridge, 1885b: 94 [urn:lsid:nmbe.ch:spidergen:06048] (type B. benevola O. Pickard-Cambridge, 1885) by Simon, 1898a: 362 and of Evippella Strand, 1906b: 680 [urn:lsid:nmbe.ch:spidergen:05800] (type E. typica Strand, 1906) by Alderweireldt, 1991: 360.

Transferred to other genera:

Evippa apsheronica Marusik, Guseinov & Koponen, 2003 -- see Halocosa

Evippa differta Lawrence, 1952 -- see Evippomma

Evippa eberlanzi (Roewer, 1959) -- see Proevippa

Evippa mandlaensis Gajbe, 2004 -- see Wadicosa

Evippa ovambica (Lawrence, 1927) -- see Evippomma

Evippa peregrina (Audouin, 1826) -- see Hogna

Evippa relicta Lawrence, 1927 -- see Evippomma

Evippa sinensis (Yin, Peng, Kim & Wang, 1995) -- see Xerolycosa

Evippa squamulata Simon, 1898 -- see Evippomma

In synonymy:

Evippa brunneopicta (Loksa, 1965, T from Xerolycosa) = Evippa sjostedti Schenkel, 1936 (Yu & Song, 1988c: 113, sub E. potanini)

Evippa fujianensis Peng, Yin & Kim, 1996 = Evippa sibirica Marusik, 1995 (Marusik & Buchar, 2004: 153)

Evippa helanshanensis Peng, Yin & Kim, 1996 = Evippa sjostedti Schenkel, 1936 (Marusik & Buchar, 2004: 153)

Evippa potanini Schenkel, 1963 = Evippa sjostedti Schenkel, 1936 (Marusik, Guseinov & Koponen, 2003b: 50)

Evippa sector Alderweireldt & Jocqué, 2017 = Evippa fortis Roewer, 1955 (Zamani et al., 2017c: 69)

Evippa straeleni Roewer, 1959 = Evippa strandi Lessert, 1926 (Alderweireldt, 1991: 372)

Nomina dubia:

Evippa abacata (Karsch, 1881a: 9, j, Libya, originally in Pardosa) -- Alderweireldt, 1991: 376

Evippa africana Strand, 1906b: 679, 1908k: 365, pl. 8, f. 10-12 (mf, Ethiopia) -- Alderweireldt, 1991: 376

Evippa brevipes Strand, 1906b: 680, 1908k: 369, pl. 8, f. 15 (f, Tunisia) -- Alderweireldt, 1991: 378

Evippa typica (Strand, 1906b: 680, 1908k: 371, pl. 8, f. 14a-c, m, Tunisia, originally in Evippella) -- Alderweireldt, 1991: 378

Evippa ungulata (O. Pickard-Cambridge, 1876b: 603, j, Egypt, originally in Lycosa) -- Alderweireldt, 1991: 376

Evippa aculeata (Kroneberg, 1875) | | Central Asia urn:lsid:nmbe.ch:spidersp:017771
Lycosa aculeata Kroneberg, 1875: 38, pl. 5, f. 46 (Df) Original description
Evippa aculeata Simon, 1895c: 342
Evippa kronebergi Roewer, 1955c: 154 (superfluous replacement name for Lycosa aculeata, erroneously considered preoccupied by Sundevall, 1833a)
Evippa aequalis Alderweireldt, 1991 | | Senegal, Sudan urn:lsid:nmbe.ch:spidersp:017757
Evippa aequalis Alderweireldt, 1991: 367, f. 3.1-5 (Dmf) Original description
Evippa amitaii Armiach Steinpress, Alderweireldt, Cohen, Chipman & Gavish-Regev, 2021 | | Israel, Iraq, Iran urn:lsid:nmbe.ch:spidersp:052803
Evippa amitaii Armiach Steinpress et al., 2021: 104, f. 4A, 5A, 6A, 7A, 8A, 9, 10A, 11A, 12A-D, 13A, C (Dmf) Original description
Evippa amitaii Zamani et al., 2022b: 368, f. 37-55 (mf)
Evippa amitaii Zamani & Marusik, 2022c: 298, f. 6A-C (m)
Evippa amitaii Al-Khazali et al., 2023: 107, f. 9A-D, 10A-J (mf)
Evippa amitaii Al-Yacoub & Najim, 2023: 385, f. 3 (f)
Evippa arenaria (Audouin, 1826) | | Algeria, Tunisia, Libya, Chad, Egypt, Israel, Palestine urn:lsid:nmbe.ch:spidersp:017758
Lycosa arenaria Audouin, 1826: 367, pl. 4, f. 3 (Df) Original description
Lycosa festiva Pavesi, 1880b: 369 (Df)
Evippa arenaria Simon, 1898a: 354, f. 346-347, 350
Evippa arenaria Roewer, 1959b: 175, f. 90a, 91 (f; misidentified per Armiach Steinpress et al., 2021: 107)
Evippa arenaria Denis, 1966e: 127, f. 41 (f; misidentified per Armiach Steinpress et al., 2021: 107)
Evippa arenaria Alderweireldt, 1991: 363, f. 1.1-3, 2.1-6 (f, Dm)
Evippa arenaria Armiach Steinpress et al., 2021: 107, f. 3A, 4B, 5B, 6B, 7B, 8B, 10B, 11B (mf)
Evippa badchysica Sternbergs, 1979 | | Turkmenistan urn:lsid:nmbe.ch:spidersp:017759
Evippa badchysica Sternbergs, 1979b: 65, f. 1 (Df) Original description
Evippa badchysica Marusik, Guseinov & Koponen, 2003b: 48, f. 23-24 (f)
Evippa baltoroi (Caporiacco, 1935) | | India (Kashmir), Nepal?, China? urn:lsid:nmbe.ch:spidersp:017192
Pardosa baltoroi Caporiacco, 1935b: 233, pl. 5, f. 17 (Dmf; N.B.: baltistana in legend) Original description
Acantholycosa baltoroi Roewer, 1955c: 152
Acantholycosa baltoroi Buchar, 1976: 202, f. 1-3 (mf; misidentified per Sankaran & Caleb, 2023b: 539)
Acantholycosa baltoroi Chen, Song & Kim, 1998: 72, f. 13-19 (mf; misidentified per Sankaran & Caleb, 2023b: 539)
Acantholycosa baltoroi Song, Zhu & Chen, 1999: 310, f. 186A, M (mf; misidentified per Sankaran & Caleb, 2023b: 539)
Acantholycosa baltoroi Song, Zhu & Chen, 2001: 226, f. 138A-D (mf; misidentified per Sankaran & Caleb, 2023b: 539)
Acantholycosa baltoroi Marusik, Azarkina & Koponen, 2004: 112, f. 60 (m; misidentified per Sankaran & Caleb, 2023b: 539)
Acantholycosa baltoroi Marusik & Omelko, 2017a: 597, f. 10 (m; misidentified per Sankaran & Caleb, 2023b: 539)
Acantholycosa baltoroi Zhang, Peng & Zhang, 2022: 119, f. 83A-J (mf; misidentified per Sankaran & Caleb, 2023b: 539)
Evippa baltoroi Sankaran & Caleb, 2023b: 539, f. 4A-B, 5A-D (mf, T from Acantholycosa)
Evippa banarensis Tikader & Malhotra, 1980 | | India urn:lsid:nmbe.ch:spidersp:017760
Evippa banarensis Tikader & Malhotra, 1980: 305, f. 121-125 (Dmf) Original description
Evippa banarensis Sankaran, Caleb & Sebastian, 2021d: 172 (species inquirenda, probably S of E. rajasthanea)
Evippa benevola (O. Pickard-Cambridge, 1885) | | China (Yarkand), India? urn:lsid:nmbe.ch:spidersp:017761
Boebe benevola O. Pickard-Cambridge, 1885b: 95 (Df) Original description
Evippa benevola Simon, 1898a: 360
Evippa beschkentica Andreeva, 1976 | | Central Asia urn:lsid:nmbe.ch:spidersp:017762
Evippa beschkentica Andreeva, 1976: 48, f. 56-58 (Dmf) Original description
Evippa brevicymbium Alderweireldt & Jocqué, 2017 | | United Arab Emirates urn:lsid:nmbe.ch:spidersp:049315
Evippa brevicymbium Alderweireldt & Jocqué, 2017: 10, pl. 6-8,12, f. 6-7, 12, 15 (Dmf) Original description
Evippa caucasica Marusik, Guseinov & Koponen, 2003 | | Azerbaijan urn:lsid:nmbe.ch:spidersp:038680
Evippa caucasica Marusik, Guseinov & Koponen, 2003b: 52, f. 4-6 (Dm) Original description
Evippa caucasica Azarkina & Nuruyeva, 2021: 340, f. 1-11 (m, Df)
Evippa concolor (Kroneberg, 1875) | | Tajikistan urn:lsid:nmbe.ch:spidersp:017763
Lycosa concolor Kroneberg, 1875: 38, pl. 4, f. 26 (Dm) Original description
Evippa concolor Simon, 1895c: 342
Evippa douglasi Hogg, 1912 | | China urn:lsid:nmbe.ch:spidersp:017764
Evippa douglasi Hogg, 1912: 215, f. 8 (Df) Original description
Evippa eltonica Dunin, 1994 | | Russia (Europe), Kazakhstan, Iran urn:lsid:nmbe.ch:spidersp:017765
Evippa eltonica Dunin, 1994b: 243, f. 1-4 (Dmf) Original description
Evippa eltonica Marusik, Guseinov & Koponen, 2003b: 50, f. 7-9, 13-15 (mf)
Evippa eltonica Esyunin, Tuneva & Farzalieva, 2007: 51, f. 6-9 (mf)
Evippa eltonica Shafaie et al., 2022: 238, f. 21-29 (m)
Evippa fortis Roewer, 1955 | | Iran, United Arab Emirates urn:lsid:nmbe.ch:spidersp:017766
Evippa fortis Roewer, 1955a: 758, f. 4 (Dm) Original description
Evippa sector Alderweireldt & Jocqué, 2017: 13, pl. 9-11, 14, f. 10-11, 14, 17 (Dmf) [urn:lsid:nmbe.ch:spidersp:049316]
Evippa fortis Zamani et al., 2017c: 69, f. 7A-C (m, S of Evippa sector)
Evippa jabalpurensis Gajbe, 2004 | | India urn:lsid:nmbe.ch:spidersp:040074
Evippa jabalpurensis Gajbe, 2004c: 10, f. 10-13 (Df) Original description
Evippa jabalpurensis Sankaran, Caleb & Sebastian, 2021d: 160, f. 1A-C (f)
Evippa jocquei Alderweireldt, 1991 | | North Africa, United Arab Emirates urn:lsid:nmbe.ch:spidersp:017769
Evippa jocquei Alderweireldt, 1991: 374, f. 8.1-3, 9 (Dmf) Original description
Evippa jocquei Alderweireldt & Jocqué, 2017: 15, pl. 8-9, 13, f. 16 (mf)
Evippa jocquei Azarkina & Trilikauskas, 2019: 556, f. 11-14, 19-25 (mf)
Evippa kazachstanica Ponomarev, 2007 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:041384
Evippa kazachstanica Ponomarev, 2007b: 94, f. 32 (Df) Original description
Evippa kirchshoferae Roewer, 1959 | | Tunisia, Spain urn:lsid:nmbe.ch:spidersp:017770
Evippa kirchshoferae Roewer, 1959b: 178, f. 121 (Df) Original description
Evippa kirschoferae Alderweireldt, 1991: 371, f. 6.1 (f)
Evippa kirchshoferae Barrientos, Miñano & Mestre, 2015: 4, f. 2A-D, 3, 4, 5A-C, 6A-B (f, Dm)
Evippa lugubris Chen, Song & Kim, 1998 | | China urn:lsid:nmbe.ch:spidersp:017772
Evippa lugubris Chen, Song & Kim, 1998: 70, f. 1-6 (Dm) Original description
Evippa lugubris Song, Zhu & Chen, 1999: 320, f. 202I (m)
Evippa luteipalpis Roewer, 1955 | | Iran urn:lsid:nmbe.ch:spidersp:017773
Evippa luteipalpis Roewer, 1955a: 759, f. 5-7 (Dmf) Original description
Evippa luteipalpis Zamani et al., 2022b: 370, f. 56-59 (mf)
Evippa massaica (Roewer, 1959) | | Tanzania urn:lsid:nmbe.ch:spidersp:017774
Evippella massaica Roewer, 1959b: 195, f. 103a-b (Df) Original description
Evippa massaica Alderweireldt, 1991: 379 (misplaced in Evippa, affinities to Pardosa)
Evippa nigerrima (Miller & Buchar, 1972) | | Afghanistan urn:lsid:nmbe.ch:spidersp:017775
Evippella nigerrima Miller & Buchar, 1972: 394, f. 26-30 (Df) Original description
Evippa onager Simon, 1895 | | Israel, Iran, Turkmenistan, China urn:lsid:nmbe.ch:spidersp:017776
Evippa onager Simon, 1895c: 341 (Df) Original description
Evippa ?onager Sternbergs, 1979b: 67, f. 1 (f, Dm; misidentified per Marusik, Guseinov & Koponen, 2003b: 50)
Evippa ?onager sensu Šternbergs 1979 Marusik, Guseinov & Koponen, 2003b: 50, f. 19-22, 28-29 (mf)
Evippa caucasica Zamani et al., 2016: 107, f. 28-30 (m) [misidentified per Zamani et al., 2017c: 63, as 'E. onager sensu Sternbergs, 1979']
Evippa onager sensu Šternbergs 1979 Armiach Steinpress et al., 2021: 111, f. 3C-D, 4C, 5C, 6C, 7C, 8C, 10C, 11C (mf)
Evippa praelongipes (O. Pickard-Cambridge, 1871) | | Libya to Saudi Arabia, Israel, Kazakhstan, Turkmenistan?, Pakistan?, India? urn:lsid:nmbe.ch:spidersp:017778
Lycosa praelongipes O. Pickard-Cambridge, 1871c: 822, pl. 50, f. 3 (Df) Original description
Pardosa praelongipes Schmidt, 1895: 476 (presumably misidentified per Armiach Steinpress et al., 2021: 113)
Evippa praelongipes Simon, 1897i: 290
Evippa praelongipes Roewer, 1959b: 182, f. 93 (f)
Evippa praelongipes Denis, 1966e: 127, f. 44 (f)
Evippa praelongipes Tikader & Malhotra, 1980: 311, f. 135-137 (f; misidentified per Armiach Steinpress et al., 2021: 113; probably to E. rajasthanea per Sankaran, Caleb & Sebastian, 2021d: 172)
Evippa praelongipes Alderweireldt, 1991: 369, f. 5.1-5 (f, Dm)
Evippa praelongipes Armiach Steinpress et al., 2021: 113, f. 3B, 4D, 5D, 6D, 7D, 8D, 10D, 11D, 14A (mf)
Evippa projecta Alderweireldt, 1991 | | Kenya urn:lsid:nmbe.ch:spidersp:017779
Evippa projecta Alderweireldt, 1991: 368, f. 4.1-3 (Dm) Original description
Evippa rajasthanea Tikader & Malhotra, 1980 | | India urn:lsid:nmbe.ch:spidersp:017780
Evippa rajasthaneus Tikader & Malhotra, 1980: 306, f. 126-130 (Dmf) Original description
Evippa rajasthanea Sankaran, Caleb & Sebastian, 2021d: 160, f. 2A-C, 3A-F (mf)
Evippa rubiginosa Simon, 1885 | | India urn:lsid:nmbe.ch:spidersp:017781
Evippa rubiginosa Simon, 1885h: 11 (Df) Original description
Evippa rubiginosa Gravely, 1924: 611, f. 5E (f)
Evippa rubiginosa Tikader & Malhotra, 1980: 313, f. 143-145 (f)
Evippa rubiginosa Sankaran, Caleb & Sebastian, 2021d: 172 (species inquirenda)
Evippa russellsmithi Alderweireldt, 1991 | | Ethiopia, Somalia urn:lsid:nmbe.ch:spidersp:017782
Evippa russellsmithi Alderweireldt, 1991: 373, f. 7.1-5 (Dmf) Original description
Evippa schenkeli Sternbergs, 1979 | | Turkmenistan urn:lsid:nmbe.ch:spidersp:017783
Evippa schenkeli Sternbergs, 1979b: 65, f. 1 (Df) Original description
Evippa schenkeli Marusik, Guseinov & Koponen, 2003b: 50, f. 25-27 (f)
Evippa shivajii Tikader & Malhotra, 1980 | | India urn:lsid:nmbe.ch:spidersp:017784
Evippa shivajii Tikader & Malhotra, 1980: 312, f. 138-142 (Dmf) Original description
Evippa shivajii Sankaran, Caleb & Sebastian, 2021d: 164, f. 4A-C, 5A-D, 6A-C (mf)
Evippa shivajii Dave & Trivedi, 2024: 24933, f. 16 (f)
Evippa sibirica Marusik, 1995 | | Kazakhstan, Russia (South Siberia), Mongolia, China urn:lsid:nmbe.ch:spidersp:017785
Evippa sibirica Marusik in Eskov & Marusik, 1995: 64, f. 58-62 (Dmf) Original description
Evippa fujianensis Peng, Yin & Kim, 1996b: 71, f. 1-7 (Dmf) [urn:lsid:nmbe.ch:spidersp:017767]
Evippa fujianensis Yin et al., 1997c: 7, f. 1a-g (mf)
Evippa fujianensis Song, Zhu & Chen, 1999: 320, f. 190I, T (mf)
Evippa sibirica Marusik, Guseinov & Koponen, 2003b: 50, f. 10-12, 16-18 (mf)
Evippa sibirica Marusik & Buchar, 2004: 153 (S of Evippa fujianensis)
Evippa sjostedti Schenkel, 1936 | | Georgia, Kazakhstan, Central Asia, Mongolia, China urn:lsid:nmbe.ch:spidersp:017786
Evippa sjöstedti Schenkel, 1936b: 304, f. 106 (Df) Original description
Evippa potanini Schenkel, 1963: 387, f. 224 (Df) [urn:lsid:nmbe.ch:spidersp:017777]
Xerolycosa brunneopicta Loksa, 1965: 16, f. 24 (Df) [urn:lsid:nmbe.ch:spidersp:056315]
Evippa potanini Song, Yu & Yang, 1982: 209, f. 1-3 (f, Dm)
Evippa potanini Hu, 1984: 216, f. 226.1-4 (mf)
Evippa potanini Song, 1987: 220, f. 178 (mf)
Evippa potanini Yu & Song, 1988c: 113 (S of Xerolycosa brunneopicta)
Evippa potanini Hu & Wu, 1989: 201, f. 167.1-4 (mf)
Evippa potanini Zhao, 1993: 67, f. 27a-c (mf)
Evippa potanini Zhao, 1995: 786, f. 363a-c (mf)
Evippa helanshanensis Peng, Yin & Kim, 1996b: 72, f. 8-11 (Df) [urn:lsid:nmbe.ch:spidersp:017768]
Evippa helanensis Yin et al., 1997c: 8, f. 2a-d (f, lapsus)
Evippa helanshanensis Song, Zhu & Chen, 1999: 320, f. 190J (f)
Evippa potanini Song, Zhu & Chen, 1999: 320, f. 190K, 191A (mf)
Evippa potanini Hu, 2001: 171, f. 76.1-4 (mf)
Evippa sjostedti Marusik, Guseinov & Koponen, 2003b: 50, f. 30-34 (mf, S of Evippa potanini)
Evippa sjostedti Marusik & Buchar, 2004: 153 (S of Evippa helanshanensis)
Evippa sjostedti Zhu & Zhang, 2011: 262, f. 189A-E (mf)
Evippa sjostedti Zhang, Peng & Zhang, 2022: 130, f. 94A-H (mf)
Evippa sjostedti Esyunin & Kabdrakhimov, 2023: 22, f. 2A (f)
Evippa sjostedti Seropian et al., 2024: 103, f. 23-26 (mf)
Evippa soderbomi Schenkel, 1936 | | Mongolia, China urn:lsid:nmbe.ch:spidersp:017787
Evippa söderbomi Schenkel, 1936b: 302, f. 105 (Dmf) Original description
Evippa soderbomi Hu & Wu, 1989: 202, f. 168.1-4 (mf)
Evippa soderbomi Song, Zhu & Chen, 1999: 320, f. 190L, 191B (mf)
Evippa sohani Tikader & Malhotra, 1980 | | India urn:lsid:nmbe.ch:spidersp:017788
Evippa sohani Tikader & Malhotra, 1980: 303, f. 116-120 (Dmf) Original description
Evippa sohani Sankaran, Caleb & Sebastian, 2021d: 166, f. 7A-B, 8A-F (mf)
Evippa solanensis Tikader & Malhotra, 1980 | | India urn:lsid:nmbe.ch:spidersp:017789
Evippa solanensis Tikader & Malhotra, 1980: 308, f. 131-134 (Df) Original description
Evippa solanensis Sankaran, Caleb & Sebastian, 2021d: 169, f. 9A-C (f)
Evippa strandi (Lessert, 1926) | | Congo, Rwanda, Tanzania urn:lsid:nmbe.ch:spidersp:017790
Proevippa strandi Lessert, 1926: 352, f. 13 (Df) Original description
Evippa straeleni Roewer, 1959b: 185, f. 96a-b (Df) [urn:lsid:nmbe.ch:spidersp:056316]
Proevippa strandi Roewer, 1959b: 193, f. 90b, 100a-b (f)
Evippa strandi Alderweireldt, 1991: 372, f. 6.3 (f, T from Proevippa, S of Evippa straeleni)
Evippa turkmenica Sternbergs, 1979 | | Kazakhstan, Turkmenistan urn:lsid:nmbe.ch:spidersp:017791
Evippa turkmenica Sternbergs, 1979b: 67, f. 1 (Df) Original description
Evippa turkmenica Marusik, Guseinov & Koponen, 2003b: 50, f. 35 (f)

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